Migration
of Birds
Influence of Topography
The relation of the world's land masses to each
other and the distribution of ecological
communities within these land masses influence
the direction birds migrate. Topography may aid,
hinder, or prevent the progress of a migrant
depending on the bird's particular requirements.
Old World migrants must contend with east and
west trending mountain ranges and deserts, and
major migratory routes tend to be in a northeast
to southwest or a northwest to southeast
direction in order to circumnavigate these
barriers. In the New World, however, migrants can
proceed north and south across the land
unimpeded, since the major mountain ranges and
river systems are oriented in the same direction
as the birds' migration.
Distinct features in the landscape, including
rivers, mountain ridges, desert rims, or
peninsulas appear to influence migratory travel
by providing a landscape reference for
orientation, especially when it is necessary to
compensate for wind drift.
Large bodies of water constitute real barriers to
soaring birds dependent on thermals, since water
temperatures are usually less than land
temperatures during both vernal and autumnal
migratory periods and thus are characterized by
subsidence of air rather than updrafts. The
shoreline, then, may provide a guiding line,
since onshore winds rise upward once they move
across the warmer land. These conditions often
concentrate Broad-winged, Rough-legged,
Red-shouldered, and Red-tailed hawks migrating
through the Great Lakes into restricted areas
where numbers observed can be spectacular. It has
been observed around Lake Ontario, for example,
that maximum hawk flights occur when winds are
from 10 to 25 miles per hour, but when winds
exceed 35 miles per hour good soaring conditions
are curtailed and hawk migration ceases. Similar
conditions exist over the Bosphorus between the
Black Sea and the Mediterranean where thousands
of White Storks, eagles, and buzzards can be
observed on a good day. For migrants not
dependent upon soaring flight, on the other hand,
large bodies of water do not affect their rate of
migration or the routes they choose. The Gulf of
Mexico, the Mediterranean Sea, and even the open
Atlantic from the Maritime Provinces of Canada to
the northern coast of South America are regularly
crossed by many songbirds.
As previously noted, mountain ridges that
parallel the line of flight offer updrafts to
soaring birds. The highest and longest ridges
deflect the horizontal winds upward better than
the shorter ridges less than 1,000 feet high, and
more birds are seen, on the average, along these
higher ridges.
In summary, topography may help or deter a
migrant in its passage. It affects different
birds in different ways. In North America,
migratory movements are continent wide, and no
evidence indicates any particular part of the
landscape influences all birds in the same
manner. Certain bird populations may use regular
geographic routes during migration, but they are
usually not rigidly restricted to them because of
topography.
Migration is dangerous. Untold thousands of
smaller migrants die each year from storms and
attacks by predators. Indeed, the passage of
migrants is so reliable that Eleonora's Falcon
breeds in the fall to take advantage of the many
songbirds crossing the Mediterranean as a source
of food for its young. Mortality during migratory
flight, of course, is one of the several costs
that balance the increased production of
offspring that migrants obtain by nesting in
locations where food is more abundant and
interspecific competition for most resources is
lower.
Storms
Of all the hazards confronting birds in
migration, storms are one of the most dangerous.
Birds that cross broad stretches of water can
confront headwinds associated with a storm,
become exhausted, and fall into the waves. Such a
catastrophe was once seen from the deck of a
vessel in the Gulf of Mexico, 30 miles off the
mouth of the Mississippi River. Great numbers of
migrating birds, chiefly warblers, were nearing
land after having accomplished nearly 95 percent
of their long flight when, caught by a
"norther" against which they were
unable to make headway, hundreds were forced into
the waters of the Gulf and drowned. A sudden drop
in temperature accompanied by a snowfall can
cause a similar effect.
Aerial Obstructions
Lighthouses, tall buildings, monuments,
television towers, and other aerial obstructions
have been responsible for destruction of
migratory birds. Bright beams of lights on
buildings and airport ceilometers have a powerful
attraction for nocturnal air travelers that may
be likened to the fascination for lights
exhibited by many insects, particularly
night-flying moths. The attraction is most
noticeable on foggy nights when the rays have a
dazzling effect that not only lures the birds but
confuses them and causes their death by collision
against high structures. The fixed, white,
stationary light located 180 feet above sea level
at Ponce de Leon Inlet (formerly Mosquito Inlet),
Florida, has caused great destruction of bird
life even though the lens is shielded by wire
netting. Two other lighthouses at the southern
end of Florida, Sombrero Key and Fowey Rocks,
have been the cause of a great number of bird
tragedies, while heavy mortality has been noted
also at some of the lights on the Great Lakes and
on the coast of Quebec. Fixed white lights seem
to be most attractive to birds; lighthouses
equipped with flashing or red lights do not have
the same attraction.
For many years in Washington, D.C., the
illuminated Washington Monument, towering more
than 555 feet into the air, caused destruction of
large numbers of small birds. Batteries of
brilliant floodlights grouped on all four sides
about the base illuminate the Monument so
brilliantly, airplane pilots noticed that it
could be seen for 40 miles on a clear night. On
dark nights with gusty, northerly winds,
nocturnal migrants seem to fly at lower altitudes
and are attracted to the Monument. As they mill
about the shaft, they are dashed against it by
eddies of wind, and hundreds have been killed in
a single night.
In September 1948, bird students were startled by
news of the wholesale destruction of Common
Yellowthroats, American Redstarts, Ovenbirds, and
others against the 1,250 foot Empire State
Building in New York City, the 491 foot
Philadelphia Saving Fund Society Building in
Philadelphia, and the 450 foot WBAL radio tower
in Baltimore. In New York, the birds continued to
crash into the Empire State Building for 6 hours.
More recently, television towers have become a
major hazard. These structures are so tall,
sometimes over 1,000 feet, they present a greater
menace than buildings or lighthouses. Their
blinking lights cause passing migrants to blunder
into guy wires or the tower itself. Numerous
instances throughout the United States indicate
this peril to migration is widespread. Yet TV
tower kills have been an excellent source of
scientific information on the fat loads migrants
carry, since they literally remove birds from out
of the sky during their migration.
Exhaustion
The American Golden-Plover travels over a
2,400-mile oceanic route from Nova Scotia to
South America in about 48 hours of continuous
flight. This is accomplished with the consumption
of less than 2 ounces of body fat. In contrast,
to be just as efficient in operation, a
1,000-pound airplane would consume only a single
pint of fuel in a 20-mile flight rather than the
gallon usually required. Similarly, the tiny
Ruby-throated Hummingbird weighing approximately
4 grams, crosses the Gulf of Mexico in a single
flight of more than 500 miles while consuming
less than 1 gram of fat.
One might expect the exertion required for long
migratory flights would result in arrival of
migrants at their destination near a state of
exhaustion. This is usually not the case. Birds
that have recently arrived from a protracted
flight over land or sea sometimes show evidences
of being tired, but their condition is far from
being in a state of exhaustion, unless they have
faced adverse winds. In reality, even small
landbirds are so little exhausted by ocean
voyages, they not only cross the Gulf of Mexico
at its widest point but may even proceed without
pause many miles inland before stopping. The
Sora, considered such a weak flyer that at least
one writer was led to infer most of its migration
was made on foot, has one of the longest
migration routes of any member of the rail family
and even crosses the wide reaches of the
Caribbean Sea. Observations indicate that under
favorable conditions birds can fly when and where
they please and the distance covered in a single
flight is governed chiefly by the rate of
dehydration and to a lesser degree, the amount of
stored fat.
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