Origin of
Bird Flight
Exactly how birds acquired the ability to fly has
baffled scientists for years. Archaeopteryx
provided a starting point for speculation. Built
like a dinosaur, but with wings, scientists
guessed at how a hypothetical ancestor might have
taken flight. Some scientists support the
arboreal hypothesis and suggest that the
ancestors of Archaeopteryx lived in
trees and glided into flapping flight. But others
argue that the claws of Archaeopteryx
weren't suited to climbing. So, others support
the cursorial hypothesis and suggest that these
ancestors used their long, powerful legs to run
fast with their arms outstretched, and were at
some point lifted up by air currents and carried
into flapping flight.
Studying living animals can throw light on their
evolutionary past. Ken Dial of the Flight Lab at the University
of Montana noticed the ability of gamebird
chicks to escape danger by scrambling up vertical
surfaces. The chicks first run very fast,
flapping their immature, partially feathered
wings, frantically creating enough momentum to
run up a vertical surface to safety. Could this
survival instinct be the origin of flight?
Bird Flight
Flight requires lift, which results when an air
stream passing over wing must travel further (and
faster) than the air stream passing under a wing.
The slower moving air under the wing 'pushes'
against the bottom of the wing with greater force
than the faster moving air above the wing &
this generates lift. The lift generated is
influenced by a bird's velocity
Why does the slower moving air generate more
pressure against the wing than the faster moving
air? In calm air, the molecules are moving
randomly in all directions. However, when air
begins to move, most (but not all) molecules are
moving in the same direction. The faster the air
moves, the greater the number of air molecules
moving in the same direction. So, air moving a
bit slower will have more molecules moving in
other directions. In the case of a wing, because
air under the wing is moving a bit slower than
air over the wing, more air molecules will be
striking the bottom of the wing than will be
striking the top of the wing. This is called the Bernoulli
effect &
this creates lift!
Wings also provide lift through Newton's
Third Law of Motion which states that for every action
there is an equal and opposite reaction. As the
wing moves though the air, the lower surface of
the wing deflects some of the air downward. As
Newton's Third Law of Motion explains, an
additional force is generated. The deflected
airflow underneath the wing is the action. The
reaction is that the wing moves in the opposite
direction (in this case, upwards). This means
that the development of low pressure above the
wing (Bernoulli's
Principle)
and the wing's reaction to the deflected air
underneath it (Newton's third Law) both
contribute to the total lift force generated.
When the curvature over the top becomes greater
by increasing the angle of attack), the air moves
even faster over the top of the wing and more
lift is generated. Eventually, however, if the
angle of attack becomes too great, the flow
separates off the wing and less lift is
generated. The result is stalling. For birds, the
optimum angle of attack is typically about 3 - 5
degrees. Birds also tend to stall at low speeds
because slower moving air may not move smoothly
over the wing.
Of course, a wing moving through the air is
opposed by friction & this is called drag.
The two main types of drag acting on birds are
pressure (or induced) drag & friction (or
profile) drag. Induced drag occurs when the air
flow separates from the surface of a wing, while
friction drag is due to the friction between the
air and bird moving through the air. Friction
drag is minimized by a wing's thin leading edge
(wings 'slice' through the air). Induced drag
occurs at low speeds and at higher speeds as, at
wing tips, air moves from the area of high
pressure (under the wing) to the area of low
pressure (top of the wing). As wings move through
the air, this curling action causes spirals
(vortices) of air which can disrupt the smooth
flow of air over a wing (and reduce lift). The
amount of drag varies with a bird's mass
(increased mass = increased friction drag) &
speed (increased speed = increased induced drag
at the wing tips), and with a wing's surface area
& shape. As described below, some wing shapes
help to reduce induced drag. Wing shapes vary
substantially among birds.
A convenient way to describe the shape of a wing
is by its aspect ratio - the ratio of length to
width. Among bird wings, aspect ratios vary from
about 1.5 to as high as about 18. Elliptical (or
'rapid takeoff') wings have relatively low aspect
ratios, while high speed wings & soaring
wings have high aspect ratios.
The long (or soaring) wings of birds with very
high aspect ratios, like albatrosses, generate
lots of lift, while the narrow, pointed shape
helps reduce drag while gliding (because the
small area of the pointed tip minimizes pressure
differences and, therefore, turbulence at the
wing tip).
High speed wings, like those of falcons, swallows
& swifts, have relatively high aspect ratios.
These narrow, tapering wings can be flapped
rapidly to generate lots of speed with minimal
drag (because, again, the small area of the
pointed tip minimizes pressure differences and
turbulence at the wing tip).
High-lift wings have lower aspect ratios &
there are spaces between the feathers at the end
of the wing. These 'slots' help reduce drag at
slow speeds because the separated tip feathers
act as 'winglets' and spread vorticity both
horizontally and vertically.
Wings with low aspect ratios (elliptical wings),
like those of many songbirds, woodpeckers,
pheasants & quail, permit sharp turns while
flying among trees & shrubs.
Another important factor that influences a bird's
flying ability is wing loading - the weight (or
mass) of a bird divided by wing area (grams/total
wing area in square centimeters). Birds with low
wing loading need less power to sustain flight.
Birds considered to be the 'best' flyers, such as
swallows & swifts, have lower wing loading
values than other birds.
| Species
or group |
|
Wing-loading |
| Swallows |
|
0.15 |
| Passerines |
|
0.2 - 0.4 |
| Hawks |
|
0.3 - 0.5 |
| Waterfowl
|
|
0.8 - 1.0 |
| Pied-billed
Grebe |
|
1.2 |
| Loons |
|
1.4 |
The high
wing loading of birds like grebes, loons, and
swans means that it's more difficult for them to
generate sufficient lift to take-off. That's why
these birds often run along the surface of a lake
for some distance before taking flight. They must
generate enough speed to generate enough lift to
get their relatively heavy bodies into the air!
Landing
Birds must usually be much more precise when
landing than an airplane pilot; often landing on
a branch rather than a runway. During landing,
birds increase the angle of attack of their wings
until they stall. This decreases both speed and
lift. Birds also spread and lower their tails,
with the tail increasing drag & acting like a
brake. Finally, legs and feet are extended for
landing.
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